The hymenium is the tissue layer on the hymenophore of a fungalfruiting body where the cells develop into basidia or asci, which produce spores. In some species all of the cells of the hymenium develop into basidia or asci, while in others some cells develop into sterile cells called cystidia (basidiomycetes) or paraphyses (ascomycetes). Cystidia are often important for microscopic identification. The subhymenium consists of the supportive hyphae from which the cells of the hymenium grow, beneath which is the hymenophoral trama, the hyphae that make up the mass of the hymenophore.
The position of the hymenium is traditionally the first characteristic used in the classification and identification of mushrooms. Below are some examples of the diverse types which exist among the macroscopic Basidiomycota and Ascomycota.
In agarics, the hymenium is on the vertical faces of the gills.
In boletes and polypores, it is in a spongy mass of downward-pointing tubes.
Gills of the fly agaric Amanita muscaria
The bolete slippery jack (Suillus luteus) with bright yellow pores
Giant puffball Calvatia gigantea
The cup fungus Sarcoscypha austriaca
The large angular pores of Polyporus alveolaris, the hexagonal-pored polypore
